Marsileaceae

(Water-clover family)

 

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GENERA (3) include Marsilea, Pilularia, and Regnellidium.

THE MARSILEACEAE are characterized by sporocarps, which bear two to many sori. The sporocarps are firm, and compressed-ovate to globose. They may be slender-pedunculate or subsessile.

SPORANGIA are indehiscent, and short- to long-stalked. They lack an annulus.

SPORES are heterosporous, and lack chlorophyll. They are trilete. Microspores are minute, and borne in groups of 16-64 within microsporangia. Megaspores are larger, and solitary within each megasporangium.

STEMS are short- to long-creeping. Slender and often branched, they are hardly indurate but bear trichomes.

PETIOLES lack stipules.

LEAVES are erect and alternate, in two rows. Circinate in bud, they are filiform, or have a long petiole with 2 or 4 leaflets. They range from 1-40 cm in length.

THE VASCULAR CYLINDER is solenostelic.

GAMETOPHYTES are minute, and non-photosynthetic. The female gametophyte is largely contained within the megasporangium; the microgametophyte is found within the microspore.

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